Myeloid Neoplasm Classification Reference

WHO Classification of Haematolymphoid Tumours, 5th Edition (Khoury et al., Leukemia 2022) & ICC 2022 (Arber et al., Blood 2022) · Source-verified

MPN Mastocytosis MLN-TK MDS MDS/MPN AML / Acute Histiocytic/DC
Myeloid Precursor Lesions — WHO 5 (new category; formally codified)
Clonal HaematopoiesisVAF ≥2% somatic mutation · no cytopenia/neoplasm = CHIP · + unexplained cytopenia = CCUS
CHIP
Clonal haematopoiesis of indeterminate potential
VAF ≥2% · no cytopenia · no haematological disorder
CCUS
Clonal cytopenia of undetermined significance
CHIP + unexplained cytopenia · does not meet MDS criteria
Myeloproliferative Neoplasms (MPN) — WHO 5 Table 1
MPN — WHO 5JMML placed under MPN in WHO 5 · CML accelerated phase ELIMINATED · CEL "NOS" qualifier removed
BCR::ABL1-positive
Chronic myeloid leukaemia
CML · BCR::ABL1+ · t(9;22)(q34;q11)
TKI-targetable
WHO 5: CP and BP only — AP formally eliminated. BP: ≥20% blasts OR extramedullary blasts OR ↑ lymphoblasts in PB/BM
BCR::ABL1-negative MPN
Polycythaemia vera
PV
JAK2 V617F ~98% / JAK2 exon 12 ~2%
Panmyelosis · pleomorphic staghorn megakaryocytes · low EPO · phlebotomy+HU/ruxolitinib
Essential thrombocythaemia
ET
JAK2/CALR/MPL
Plt ≥450 · large hyperlobated megakaryocytes loose clusters · MF-0 · CALR type 1 vs 2 must be reported
Primary myelofibrosis
PMF — pre-PMF (MF-0/1) and overt PMF (MF-2/3)
WHO 5 two-stage entity
JAK2/CALR/MPL · naked nuclei · leukoerythroblastic PB · ruxolitinib/pacritinib/momelotinib · alloSCT
Chronic neutrophilic leukaemia
CNL
CSF3R T618I >60%
WBC ≥25×10⁹/L (WHO 5 threshold) · ≥80% segmented+banded · no dysgranulopoiesis
Chronic eosinophilic leukaemia
CEL (WHO 5 removes "NOS" qualifier)
Diagnosis of exclusion
Eos ≥1.5×10⁹/L · clonal · BM dysplasia required (new WHO 5 criterion) · no MLN-TK · no AML
Juvenile myelomonocytic leukaemia
JMML — under MPN in WHO 5 (not MDS/MPN)
WHO 5 moved JMML to MPN
RAS pathway (PTPN11/NRAS/KRAS/NF1/CBL) · KMT2A rearrangements now excluded from JMML
MPN, not otherwise specified
MPN-NOS
Diagnosis of exclusion
Mastocytosis — WHO 5 Table 2
MastocytosisCD30 + any activating KIT mutation = new minor criteria · KIT D816V VAF ≥10% = B-finding · BM mastocytosis = new subtype
Cutaneous mastocytosis (skin only)
Urticaria pigmentosa / MPCM
Monomorphic or polymorphic
Diffuse cutaneous mastocytosis
Cutaneous mastocytoma
Isolated or multilocalized
Systemic mastocytosis (SM) subtypes
Bone marrow mastocytosis
BMM — NEW WHO 5 subtype
New in WHO 5
No skin lesions · no B-findings · tryptase <125 ng/mL · Hymenoptera anaphylaxis assoc.
Indolent systemic mastocytosis
ISM
No C-findings
Smoldering systemic mastocytosis
SSM — 2+ B-findings · no C-findings
Aggressive systemic mastocytosis
ASM — C-findings present
Organ damage
SM with associated haematological neoplasm
SM-AHN (WHO 5 allows lymphoid AHN; ICC renames to SM-AMN myeloid only)
Dual clonal disease
Mast cell leukaemia
MCL · ≥20% BM mast cells
Aleukemic variant possible
Mast cell sarcoma
MCS
Rare · destructive local
Myeloid/Lymphoid Neoplasms with Eosinophilia and TK Gene Fusions (MLN-TK) — WHO 5 Table 11
MLN-TK (renamed from MLN-eo in WHO 5) — supersedes all other myeloid/lymphoid diagnoses in hierarchyFISH/RT-PCR mandatory before TKI · FIP1L1::PDGFRA is cryptic on karyotype
MLN with PDGFRA rearrangement
FIP1L1::PDGFRA del(4q12) — CRYPTIC
Imatinib dramatic response
MLN with PDGFRB rearrangement
ETV6::PDGFRB t(5;12) · >30 partners
Imatinib-sensitive
MLN with FGFR1 rearrangement
ZMYM2::FGFR1 8p11 · >15 partners
Imatinib-RESISTANT · pemigatinib/futibatinib
MLN with JAK2 rearrangement
PCM1::JAK2 t(8;9) + ETV6::JAK2 + BCR::JAK2
Ruxolitinib
MLN with FLT3 rearrangement
ETV6::FLT3 t(12;13) most common
New in WHO 5FLT3 inhibitors
MLN with ETV6::ABL1
t(9;12)(q34.1;p13.2)
TKI (dasatinib)
MLN with other defined TK fusions
ETV6::FGFR2 · ETV6::LYN · ETV6::NTRK3 · RANBP2::ALK · BCR::RET · FGFR1OP::RET
Scalable landing spot
Myelodysplastic Neoplasms (MDS) — WHO 5 Table 3 (renamed "syndromes" → "neoplasms")
MDS — WHO 5 Genetic-First Classification · SLD/MLD distinction optional · MDS-h new distinct typeGenetic entities take priority regardless of blast %
MDS with defining genetic abnormalities (classified by genetics regardless of blast %)
MDS with low blasts and isolated 5q deletion
MDS-5q
Lenalidomide-responsive
<5% BM / <2% PB blasts · del(5q) alone or +1 (not −7/del7q) · monolobate megakaryocytes
MDS with low blasts and SF3B1 mutation
MDS-SF3B1 (formerly MDS-RS)
WHO 5 genetic entity
Ring sideroblasts ≥15% or ≥5% if SF3B1 mut · better prognosis · luspatercept
MDS with biallelic TP53 inactivation
MDS-biTP53
WHO 5 genetic entityWorst MDS prognosis
2 TP53 muts OR 1 mut + del17p/cnLOH · complex karyotype · supersedes MDS-5q and MDS-SF3B1
MDS morphologically defined (no defining genetic abnormality found) — WHO 5 Table 3
MDS with low blasts
MDS-LB · WHO 5 merged SLD + MLD (distinction optional)
SLD/MLD distinction optional
<5% BM / <2% PB blasts · dysplasia ≥1 lineage
MDS, hypoplastic
MDS-h — new distinct type in WHO 5
New in WHO 5
BM ≤25% cellularity · T-cell mediated · clonal · IST-responsive subset · overlaps AA/PNH
MDS with increased blasts 1
MDS-IB1 · 5–9% BM OR 2–4% PB
HMA
MDS with increased blasts 2
MDS-IB2 · 10–19% BM OR 5–19% PB OR Auer rods
Auer rod → IB2 regardless of blast %
MDS with fibrosis
MDS-F · MF-2/3 + MDS features
BM fibrosis grade ≥2
Childhood MDS — WHO 5 updated terminology (Table 4)
Childhood MDS with low blasts
cMDS-LB — WHO 5 replaces "Refractory cytopenia of childhood (RCC)"
RCC renamed in WHO 5
Childhood MDS with increased blasts
cMDS-IB · ≥5% BM or ≥2% PB blasts
More RAS mutations than cMDS-LB
Myelodysplastic/Myeloproliferative Neoplasms (MDS/MPN) — WHO 5 Table 5
MDS/MPN — WHO 5: JMML moved to MPN · aCML renamed · CMML-0 eliminated · CMML threshold loweredOverlapping MDS + MPN features
Chronic myelomonocytic leukaemia
CMML-1 / CMML-2 (CMML-0 eliminated) · MD-CMML vs MP-CMML
TET2 · SRSF2 · ASXL1
Monocytes ≥0.5×10⁹/L AND ≥10% (lowered from 1.0) · if 0.5–1.0: clonality + dysplasia required
MDS/MPN with neutrophilia
MDS/MPN-N — WHO 5 renamed aCML
aCML renamed in WHO 5
WBC ≥13 · dysgranulopoiesis · SETBP1/ASXL1 · BCR::ABL1− · eosinophils <10%
MDS/MPN with SF3B1 mutation and thrombocytosis
MDS/MPN-SF3B1-T — WHO 5 renamed from MDS/MPN-RS-T
Renamed in WHO 5
SF3B1 mut + plt ≥450 + ring sideroblasts · JAK2/CALR co-mutation possible
MDS/MPN, not otherwise specified
MDS/MPN-NOS (WHO 5 removes "unclassifiable" term)
Diagnosis of exclusion
Acute Myeloid Leukaemia — WHO 5 Table 7 (two-family framework)
AML with defining genetic abnormalities — most diagnosed at ANY blast % · exceptions: BCR::ABL1 and CEBPA require ≥20%NUP98 rearrangement is NEW WHO 5 entity · AML with mutated TP53 is NOT a separate entity in WHO 5 (it is in ICC)
APL with PML::RARA fusion ⚠ EMERGENCY
t(15;17) · WHO 5: ≥10% blasts required
START ATRA IMMEDIATELY
HLA-DR−/CD34− on flow + DIC = APL. Do NOT wait for FISH. ATRA+ATO standard
AML with RUNX1::RUNX1T1 fusion
t(8;21) · CBF-AML · any blast %
Favorable · HiDAC · GO benefit
CD19+ aberrant · salmon-pink granules · long single Auer rods
AML with CBFB::MYH11 fusion
inv(16)/t(16;16) · CBF-AML · any blast %
Favorable · HiDAC · GO benefit
CD2+ aberrant · abnormal BM eosinophils · CBFB FISH required
AML with DEK::NUP214 fusion
t(6;9) · any blast %
Poor · FLT3-ITD common co-mut
Basophilia · dysplasia · young patients
AML with RBM15::MRTFA fusion
t(1;22) · any blast % · formerly RBM15::MKL1
Infant AMKL
Infants · CD41/CD61+ · DS-like presentation
AML with BCR::ABL1 fusion
t(9;22) · WHO 5: ≥20% blasts required (exception)
WHO 5 distinct entity · TKI + chemo
Must distinguish from CML blast crisis. WHO 5 keeps 20% to avoid CML overlap
AML with KMT2A rearrangement
11q23.3 · >80 partners · any blast %
Intermediate–poor
CD33+++ very bright · NG2+ · monocytic · revumenib (menin inhibitor FDA 2024)
AML with MECOM rearrangement
inv(3)/t(3;3) · any blast %
Worst prognosis AML · −7 co-finding
Dysplastic megakaryocytes · GATA2 repression · median OS 3–5 mo
AML with NUP98 rearrangement
NUP98 11p15.4 · >30 partners · any blast %
NEW entity in WHO 5 · NOT in ICCPoor prognosis · often cryptic on karyotype
NUP98::NSD1 · NUP98::HOXA9 etc. · HOX deregulation · RNA-seq required
AML with NPM1 mutation
~30% of all AML · any blast %
Favorable (without FLT3-ITD high AR)
CD34− paradox · cup-like nuclei · NPM1c IHC · menin inhibitors emerging
AML with CEBPA mutation
In-frame bZIP domain · WHO 5: ≥20% blasts required (exception)
Favorable · mono- or bi-allelic bZIP
WHO 5 updated: monoallelic bZIP also qualifies. Germline CEBPA risk. CD7+ aberrant
AML, myelodysplasia-related (AML-MR) — WHO 5 8-gene list (Table 8) · ≥20% blasts required
AML, myelodysplasia-related
AML-MR · WHO 5 mutation list = 8 genes (NO RUNX1): ASXL1, BCOR, EZH2, SF3B1, SRSF2, STAG2, U2AF1, ZRSR2
Replaces AML-MRC · CPX-351 standard
Prior MDS/MDS-MPN OR MDS-defining cytogenetics OR any of 8 listed gene mutations · ≥20% blasts
AML with other defined genetic alterations — landing spot for new/rare entities (replaces "provisional")
AML with other defined genetic alterations
Includes AML with rare genetic fusions e.g. KAT6A, CBFA2T3::GLIS2 (RAM immunophenotype AMKL)
Scalable framework
AML defined by differentiation — WHO 5 replaces "AML NOS" · ≥20% blasts · diagnosis of exclusion
AML defined by differentiation
Minimal diff · without maturation · with maturation · basophilic · myelomonocytic · monocytic · erythroid · megakaryoblastic
Diagnosis of exclusion
WHO 5 eliminates "NOS" term entirely. AEL: ≥30% proerythroblasts + ≥80% erythroid predominance
Secondary Myeloid Neoplasms — WHO 5 (new segregated category)
Secondary MN — WHO 5 groups therapy-related + germline predisposition herePARP inhibitors added as qualifying therapy · qualifier approach (not standalone category)
Myeloid neoplasm post cytotoxic therapy
MN-pCT (WHO 5 preferred term; replaces "therapy-related") · PARP inhibitors now included as qualifying agents · methotrexate excluded
PARP inhibitors = new qualifying therapy
Myeloid neoplasms with germline predisposition
CEBPA / DDX41 / TP53 / RUNX1 / ANKRD26 / ETV6 / GATA2 / SAMD9 / SAMD9L / BM failure syndromes / RASopathies / Down syndrome
Scalable formulaic model
Myeloid proliferations associated with Down syndrome
TAM (first 6 months) · ML-DS · GATA1 exon 2/3 somatic mutation · trisomy 21
ALAL/MPAL — WHO 5 (two-family framework) · BPDCN · Histiocytic/DC Neoplasms
ALAL/MPAL with defining genetics AND immunophenotypically-defined · BPDCN criteria updated (Table 15) · RDD + ALK+ histiocytosis = new entities
ALAL with defining genetic abnormalities (WHO 5 Table 12)
MPAL with BCR::ABL1
Ph+ · TKI + ALL-type induction · alloSCT
MPAL with KMT2A rearrangement
11q23.3 · infant · poor prognosis
MPAL with ZNF384 rearrangement
B/myeloid · EP300/TCF3/TAF15/CREBBP partners
New in WHO 5
ALAL with BCL11B rearrangement
AUL + T/myeloid MPAL + AML minimal diff + ETP-ALL
New in WHO 5
ALAL with other defined genetic alterations
PHF6 mutations · PICALM::MLLT10
ALAL immunophenotypically defined
MPAL B/myeloid · MPAL T/myeloid · MPAL rare types · ALAL NOS · AUL
BPDCN — WHO 5 Table 15 (updated immunophenotypic criteria)
Blastic plasmacytoid dendritic cell neoplasm
BPDCN
CD123++ · TCF4+ · CD303+ · CD304+ · CD4+ · CD56+
WHO 5 adds CD303/CD304 as expected positive pDC markers. Tagraxofusp-erzs FDA 2018
Mature pDC proliferation assoc. with myeloid neoplasm
MPDCP — New WHO 5 entity
New in WHO 5
Low-grade clonal pDC expansion in CMML/AML context · shared mutations (RUNX1 in AML-associated)
Histiocytic/DC neoplasms — WHO 5 Table 14 (RDD + ALK+ histiocytosis added; FDCS moved to stroma-derived)
LCH · LCS
CD1a+ · CD207/Langerin+
BRAF V600E ~55%
LCH: coffee-bean nuclei · mixed inflammatory background · vemurafenib/trametinib. LCS: malignant cytology
Indeterminate dendritic cell tumour
IDCT · CD1a+/S100+/CD207−
Interdigitating dendritic cell sarcoma
IDCS · S100+ · CD1a−
Juvenile xanthogranuloma
JXG
MAP2K1/NF1 · Touton giant cells
CD163+/Factor XIIIa+/CD1a− · children · MAPK pathway
Erdheim-Chester disease
ECD
BRAF V600E ~55%
"Hairy kidney" · coated aorta · foamy macrophages · vemurafenib first-line
Rosai-Dorfman disease
RDD — NEW entity in WHO 5
New in WHO 5
S100+/CD163+/emperipolesis · KRAS/MAP2K1 mutations · sporadic/IgG4-related forms
ALK-positive histiocytosis
NEW entity in WHO 5
New in WHO 5
KIF5B::ALK most common · remarkable ALK inhibitor response · multisystem form in infants
Histiocytic sarcoma
HS
CD68+/CD163+/Lysozyme+/CD1a− · BRAF V600E ~30%
Transdifferentiation from FL (shared IGH clonality)

Source: Khoury JD, Solary E, Hochhaus A et al. WHO 5th Edition. Leukemia 2022;36:1703–1719 · Tables 1–15. Full classification: tumourclassification.iarc.who.int/chapters/63 · ⚠ APL = medical emergency — start ATRA empirically while FISH pending

ICC 2022 (Arber et al., Blood 2022) — Key differences from WHO 5: ICC retains CML accelerated phase · JMML under Pediatric disorders · introduces MDS/AML bridging category (10–19% blasts) · retains aCML name · TP53-mutated AML = standalone entity in ICC (NOT in WHO 5) · 9-gene AML-MR list (adds RUNX1) · NUP98 rearrangement NOT an AML entity in ICC · CMUS/CCMUS = new ICC concepts · MDS/MPN with i(17q) provisional subentity · SM-AHN renamed SM-AMN
ICC 2022 — MPN (Table 1 & Table 2)
MPN — ICC 2022CML AP retained in ICC · pre-PMF and overt PMF listed as two separate PMF subtypes · CEL retains NOS qualifier
Chronic myeloid leukaemia
CML — ICC retains accelerated phase (AP)
AP: 10–19% blasts + major route ACA · BP: ≥20%
ICC Table 2: AP = blasts 10–19% OR basophils ≥20% OR ACA in Ph+ cells. Major route ACA: +Ph, +8, i(17q), +19, complex, abn 3q26.2
Polycythemia vera
PV — same criteria as WHO 5
JAK2 V617F or exon 12
Essential thrombocythemia
ET — ICC Table 4: infrequent small clusters (≤6 cells) allowed; >6 cells + granulocytic proliferation = pre-PMF
PMF — early/prefibrotic stage
pre-PMF — ICC lists as separate from overt PMF (ICC Table 5)
Major criterion: mega atypia + MF<2 + hypercellular + granulocytic proliferation + ↓ erythropoiesis
PMF — overt fibrotic stage
Overt PMF — MF 2/3 + mega atypia (ICC Table 5)
Chronic neutrophilic leukaemia
CNL — ICC lowers threshold to WBC ≥13×10⁹/L if CSF3R T618I present (vs ≥25 if absent)
CSF3R T618I
Chronic eosinophilic leukaemia, NOS
CEL, NOS — ICC retains NOS qualifier; now requires BM dysplastic megakaryocytes + clonality (ICC Table 7)
ICC adds BM morphology as required criterion
MPN, unclassifiable
MPN-U (ICC Table 9)
ICC 2022 — Mastocytosis (SM-AHN → SM-AMN; CD30 added as minor criterion)
SM — ICC 2022 (Table 11): SM-AHN renamed SM-AMN · CD30 added as minor criterion · tryptase + CD117 IHC added to major criterion
All SM subtypes same as WHO 5 except SM-AHN → SM-AMN
SM-AMN = SM with an associated myeloid neoplasm (ICC restricts to myeloid; WHO 5 SM-AHN allows lymphoid). ICC Table 11: major criterion now requires tryptase/CD117 IHC confirmation of mast cells
ICC 2022 — MDS/MPN (Table 1 continued; aCML retained; JMML moved; CMUS/CCMUS new)
MDS/MPN — ICC 2022ICC retains aCML name · CMUS/CCMUS = new ICC-only concepts · i(17q) provisional subentity · JMML moved to Pediatric
Chronic myelomonocytic leukaemia
CMML — ICC Table 13: same as WHO 5 (monocytes ≥0.5×10⁹/L AND ≥10%; CMML-0 eliminated; CMML-1/2 two-tier retained)
ICC adds clonality requirement: ≥1 myeloid mutation VAF ≥10% OR abnormal cytogenetics. If no clonality: monocytes ≥1.0×10⁹/L required
Clonal monocytosis of undetermined significance
CMUS — ICC-only concept (Table 14)
ICC-only entity
Monocytes ≥0.5×10⁹/L AND ≥10% + clonality + NO BM findings of CMML · CCMUS if cytopenia present
Atypical CML
aCML — ICC retains this name (WHO 5 renamed to MDS/MPN-N) · ICC Table 15
ICC retains aCML name
WBC ≥13 · dysgranulopoiesis · eosinophils <10% · monocytes <10% · BCR::ABL1− · SETBP1+ASXL1 support diagnosis
MDS/MPN with thrombocytosis and SF3B1 mutation
MDS/MPN-T-SF3B1 (ICC Table 16) — SF3B1 VAF >10% · plt ≥450 · anemia required
MDS/MPN-RS-T, NOS
For SF3B1-WT cases with ≥15% ring sideroblasts + thrombocytosis (ICC Table 17)
ICC retains this NOS variant
MDS/MPN with i(17q)
Provisional subentity under MDS/MPN-NOS (ICC Table 19)
ICC provisional entity only
i(17q) isolated or +1 additional (not −7/del7q) · WBC ≥13 · dysgranulopoiesis · JAK2/CALR/MPL absent
MDS/MPN, NOS
MDS/MPN-NOS (ICC Table 18) · requires cytopenia + WBC ≥13 and/or plt ≥450 · clonality expected
ICC 2022 — MDS (Table 20): SLD/MLD retained · MDS/AML bridging category (10–19% blasts) new
ICC MDS — retains SLD/MLD distinction (unlike WHO 5) · introduces MDS/AML for 10–19% blasts · TP53-mutated MDS as distinct entity
Premalignant (ICC explicitly positions here)
Clonal cytopenia of undetermined significance
CCUS — somatic mut VAF ≥2% + unexplained cytopenia · no MDS criteria met
MDS with defining genetic abnormalities (ICC Table 20)
MDS with mutated SF3B1
MDS-SF3B1 — same criteria as WHO 5
MDS with del(5q)
MDS-del(5q) — same criteria
MDS with mutated TP53
ICC entity (Table 21) · multi-hit required for MDS designation
ICC-specific standalone entity
Multi-hit TP53 (2 muts or 1 mut + del17p/cnLOH/VAF>50%) for MDS. Any somatic TP53 VAF>10% sufficient for MDS/AML and AML. Part of "myeloid neoplasms with mutated TP53" spectrum
MDS, NOS — ICC retains SLD and MLD as separate subtypes (unlike WHO 5 MDS-LB)
MDS, NOS without dysplasia
−7/del7q or complex karyotype + cytopenia + no dysplasia
MDS, NOS with single lineage dysplasia
MDS-NOS SLD — retained in ICC
ICC retains SLD (WHO 5 merged into MDS-LB)
MDS, NOS with multilineage dysplasia
MDS-NOS MLD — retained in ICC
ICC retains MLD (WHO 5 merged into MDS-LB)
MDS with excess blasts
MDS-EB — single tier in ICC: 5–9% BM or 2–9% PB (no IB1/IB2 distinction like WHO 5)
ICC has only ONE MDS-EB tier
MDS/AML — ICC-only bridging category (NOT in WHO 5) · 10–19% blasts
MDS/AML — new ICC category
10–19% blasts in BM or PB · eligible for MDS AND AML trials
ICC-only · not in WHO 5
Sub-classified as: MDS/AML with mutated TP53 · MDS/AML with MR gene mutations · MDS/AML with MR cytogenetics · MDS/AML NOS
ICC 2022 — AML (Table 25: blast thresholds for each entity)
ICC AML — 9-gene MR list (adds RUNX1 vs WHO 5 8-gene) · TP53 standalone entity · NO NUP98 entity · CEBPA ≥10% blasts
APL with PML::RARA
≥10% blasts (same as WHO 5)
⚠ EMERGENCY
AML with RUNX1::RUNX1T1
≥10% blasts
CBF-AML
AML with CBFB::MYH11
≥10% blasts
CBF-AML
AML with MLLT3::KMT2A t(9;11)
≥10% blasts · ICC names t(9;11) specifically (WHO 5 uses KMT2A rearrangement)
AML with other KMT2A rearrangements
≥10% blasts
AML with DEK::NUP214
≥10% blasts
AML with MECOM rearrangement inv(3)/t(3;3)
≥10% blasts
AML with BCR::ABL1
≥20% blasts — ICC retains 20% threshold. ICC footnote: MDS/AML category NOT used for BCR::ABL1 due to CML overlap
AML with mutated NPM1
≥10% blasts
Favorable
AML with in-frame bZIP CEBPA mutations
≥10% blasts — ICC lowers threshold vs WHO 5 (≥20%)
ICC: ≥10%; WHO 5: ≥20%Favorable · bZIP required
AML and MDS/AML with mutated TP53
MDS/AML: 10–19% blasts · AML: ≥20% blasts · any somatic TP53 VAF >10%
ICC-only standalone entityNOT in WHO 5 as standalone AML entity
ICC 9-gene AML-MR mutation list (adds RUNX1 vs WHO 5 8-gene list)
AML with myelodysplasia-related gene mutations
MDS/AML (10–19%) + AML (≥20%) · ICC 9 genes: ASXL1, BCOR, EZH2, RUNX1, SF3B1, SRSF2, STAG2, U2AF1, ZRSR2
ICC adds RUNX1 vs WHO 5
AML with myelodysplasia-related cytogenetic abnormalities
MDS/AML (10–19%) + AML (≥20%) · complex karyotype + MDS-defining cytogenetics · separate from gene mutation category in ICC
AML, NOS
MDS/AML (10–19%) + AML (≥20%) · diagnosis of exclusion
ICC 2022 — Pediatric/Germline Disorders (JMML placed here, not MPN)
Pediatric — ICC places JMML here; JMML-like neoplasms + Noonan-MPD = new ICC entities; RCC retained (WHO 5 renamed to cMDS-LB)
JMML
Under Pediatric in ICC (vs MPN in WHO 5)
JMML placement differs from WHO 5
ICC Table 22: PB monocytes ≥1×10⁹/L (note: ~7% may not meet this) + splenomegaly + blasts <20% + BCR::ABL1− + RAS pathway mutation
JMML-like neoplasms
ICC-only concept (Table 23)
ICC-only
Phenotypically JMML but no RAS pathway mutation · ALK/ROS1/FIP1L1::RARA/CCDC88C::FLT3 fusions · excludes AML or MLN-TK entities
Noonan syndrome-associated myeloproliferative disorder
Transient · polyclonal · germline PTPN11/KRAS/NRAS/RIT1 · secondary mutations absent
ICC formal entity
Refractory cytopenia of childhood
RCC — ICC retains this name (WHO 5 renamed to cMDS-LB)
ICC retains RCC name
Hypocellular BM ≥80% · persistent cytopenia + dysplasia · blasts <5% BM / <2% PB · no fibrosis · germline predisposition common

Source: Arber DA, Orazi A, Hasserjian RP et al. ICC 2022. Blood 2022;140:1200–1228 · Tables 1–26. Key: ICC-only entities = CMUS/CCMUS, MDS/AML category, MDS with mutated TP53, AML with mutated TP53, JMML-like, RCC retained name, aCML retained name, MDS/MPN with i(17q), SM-AMN terminology

EntityLineageFlow Cytometry Key MarkersIHC Key MarkersCritical Molecular / Genetics
Myeloproliferative Neoplasms (MPN)
CML · BCR::ABL1+Pluripotent HSC; granulocytic CD13+CD33+CD34+ (blasts)MPO+ (granulocytes)TdT− (CP)
⚑ BC transformation: lymphoid BC = CD19+/TdT+; myeloid BC = MPO+. WHO 5 eliminates AP; ICC retains AP (10–19% blasts + major route ACA)
BCR::ABL1 FISH/RT-PCR (IS) — diagnosticMPO+CD34+ increases in AP (ICC) or BC
★ RT-PCR BCR::ABL1 International Scale (IS): aim for MMR (MR3.0); deep MR4/MR4.5 for TFR attempt. T315I = ponatinib only; Y253H/E255K = resist 1st/2nd gen TKI
t(9;22)(q34;q11) · p210 BCR::ABL1 (CML) · ASXL1 co-mut adverse in CP · Sokal/ELTS score · WHO 5: CP + BP only. ICC: CP + AP + BP. ABL1 kinase domain mutations guide TKI selection
Polycythaemia veraMultipotent HSC; erythroid CD71+CD235a (GPA)+CD36+
★ Flow limited role in PV. Gold standard: JAK2 mutation + low serum EPO + BM biopsy (panmyelosis). JAK2 exon 12: may have isolated erythrocytosis without panmyelosis
JAK2 V617F or exon 12 mutationCD71++ (erythroid hyperplasia)CD34 not increased
★ BM: panmyelosis · near-100% cellularity · fat reduced · pleomorphic megakaryocytes ("staghorn" nuclear lobation) · loose scattered clusters · BM biopsy may not be required if Hb >18.5g/dL male or >16.5g/dL female + JAK2 mutation
JAK2 V617F (~98%) · JAK2 exon 12 (~2%) · TET2 · DNMT3A · ASXL1 · LNK(SH2B3) · Post-PV MF: IDH1/2 · TP53 clonal evolution · Phlebotomy + aspirin ± HU · Ruxolitinib high-risk/refractory
ET vs pre-PMF — THE critical MPN distinctionMultipotent HSC; megakaryocytic CD41+ (GPIb)CD61+ (GPIIIa)Circulating CD34+ progenitors: <10/μL = ET · >10/μL = PMF
★ Circulating CD34+ progenitor count on PB flow = SINGLE MOST DISCRIMINATING flow finding between ET and pre-PMF when BM morphology is equivocal
Driver mutation: JAK2 V617F / CALR type 1 or 2 / MPL W515MF-0 on reticulin (ET)
★ ET megakaryocytes: large mature hyperlobated ≥5 lobes ("staghorn"), loosely scattered/small clusters · pre-PMF: TIGHT dense clusters (>6 cells), bulbous hyperchromatic nuclei, "naked nuclei" (extruded segments). CALR type MUST be reported (type 1=best OS, lower thrombosis; type 2=intermediate)
JAK2 V617F (~55%) · CALR type 1 del52bp (best OS) · CALR type 2 ins5bp · MPL W515L/K (~4%) · Triple-negative (worst outcome) · HMR co-muts: ASXL1/SRSF2/EZH2/IDH1-2/U2AF1-Q157 · ET pre-PMF distinction drives different risk/treatment approach
Primary myelofibrosis (overt)Multipotent HSC; megakaryocytic + stromal CD34+ circulating progenitors >10/μL in PBCD41+/CD61+
⚑ Teardrop cells + leukoerythroblastosis in PB = myelofibrosis until proven otherwise. Dry tap = fibrosis
Reticulin stain MF grade (0–3) mandatory in ALL MPN trephine reportsCD61+ (atypical mega clusters)
★ Overt PMF: MF-2/3 · osteosclerosis · sinusoidal hematopoiesis · naked megakaryocyte nuclei
JAK2 V617F (~55%) · CALR type 1 (best OS) · CALR type 2 · MPL (~5%) · Triple-negative (worst) · Ruxolitinib (JAK1/2) · Pacritinib (plt <50) · Momelotinib (anemia) · AlloSCT curative intent
Systemic mastocytosis (all SM subtypes)Mast cell progenitor → CD34+ BM progenitor CD117++ (c-KIT, very bright)CD25+ (ABERRANT — absent on normal mast cells)CD33+CD45+CD2+ (subset)CD30+ (new minor criterion WHO 5 + ICC)
★ CD25+/CD117++ = SM until proven otherwise. CD25 absent on reactive MCs. CD30 added as minor criterion in BOTH WHO 5 and ICC
Tryptase IHC — highlights MC aggregatesCD117++ (KIT IHC)CD25+CD30+
★ WHO SM criteria (both WHO 5 + ICC agree): Major = multifocal dense MC aggregates ≥15 MCs/cluster. 4 minor criteria (1 major + 1 minor OR 3 minor for diagnosis). WHO 5 adds BMM as new SM subtype. WDSM: round MCs, D816V absent, CD25−, CD30+ — fails standard minor criteria
KIT D816V (>95% SM) · Avapritinib (BLU-285) FDA 2021 for advanced SM/D816V · Midostaurin (broader KIT) · KIT D816V VAF ≥10% = B-finding (new WHO 5) · TET2/SRSF2/ASXL1/RUNX1 (SMAR mutations) → SM-AHN risk
MLN-TK (all subtypes)Multipotent HSC; eosinophilic predominance CD34+CD33+CD25+ (PDGFRA — MC component)
⚑ FIP1L1::PDGFRA is CRYPTIC on karyotype — FISH with PDGFRA break-apart probe or RT-PCR mandatory. NEVER diagnose CEL without excluding all MLN-TK fusions
FISH/RT-PCR: PDGFRA/B/FGFR1/JAK2/FLT3/ABL1 break-apart probes — mandatory before TKI
★ MLN-TK supersedes all other myeloid/lymphoid diagnoses (both WHO 5 + ICC). PCM1::JAK2: BM fibrosis prominent. FGFR1: most aggressive, may present as T-LBL before myeloid phase
PDGFRA: imatinib dramatic · T674I resistance (avapritinib) · PDGFRB: imatinib · FGFR1: pemigatinib/futibatinib · JAK2 (PCM1::JAK2): ruxolitinib · FLT3: midostaurin/quizartinib · ETV6::ABL1: dasatinib
Myelodysplastic Neoplasms (MDS)
MDS-5q / MDS-del(5q)Multipotent HSC; megakaryocytic + erythroid CD41+CD61+CD71+ (dysplastic erythropoiesis)
★ FISH confirmation of del(5q) required before lenalidomide. Isolated = del(5q) ± 1 additional cytogenetic abnormality (NOT −7/del7q)
Monolobate megakaryocytes on CD61 IHC — pathognomonic hallmarkCD34 not increased
★ Count nuclear lobes on CD61 IHC: del(5q) = predominantly 1 lobe (monolobate). Normal = ≥3 lobes. FISH + TP53 sequencing mandatory before lenalidomide (TP53 mutation = resistance mechanism)
del(5q) isolated or +1 (not −7/del7q) · RPS14 haploinsufficiency → macrocytic anemia · miR-145/146a loss → thrombocytosis · Lenalidomide targets CRBN · TP53 acquisition after lenalidomide = transformation risk
MDS-SF3B1 (ring sideroblast MDS)Multipotent HSC; erythroid predominance CD71+CD36+CD235a+Aberrant CD71/CD36 asynchronous maturationCD34 not increased Ring sideroblasts on Perl's Prussian blue stain (≥15% or ≥5% if SF3B1 mut)CD71++ (erythroid hyperplasia)
★ Ring sideroblast: ≥5 iron granules encircling ≥1/3 of nucleus on Perl's stain. Perl's stain MANDATORY in all MDS workups — never report without it. SF3B1 mutation causes aberrant 3' splice site → mitochondrial iron accumulation
SF3B1 splice factor mutation · Luspatercept (TGF-β trap, activin receptor ligand trap) FDA-approved for SF3B1-mutated MDS-RS · Co-mut JAK2/CALR + thrombocytosis → reclassify as MDS/MPN-SF3B1-T
MDS-biTP53 (WHO 5) / MDS with mutated TP53 (ICC)Multipotent HSC; multi-lineage dysplasia Variable blast % and myeloid phenotypeCD34+
⚑ WHO 5: biallelic TP53 required for MDS entity (2 muts OR 1 mut + del17p/cnLOH). ICC: multi-hit required for MDS; any TP53 VAF>10% sufficient for MDS/AML and AML. VAF >50% = presumptive biallelic on NGS
p53 IHC strong >10% cells = missense gain-of-function surrogatep53 IHC null = nonsense/frameshift loss-of-function
★ p53 IHC NULL does NOT exclude biallelic TP53 — molecular testing always required. Complex karyotype virtually universal. Eprenetapopt/APR-246 + aza · Magrolimab (anti-CD47)
TP53 biallelic (2 muts OR 1 mut + del17p/cnLOH) · Complex karyotype (−5, −7, −17p, +8) · Very poor OS (<12 months) · Venetoclax+aza · APR-246/eprenetapopt
MDS-LB / MDS-IB1 / MDS-IB2 / MDS-F / MDS-hMultipotent HSC; multi-lineage dysplasia CD34+: MDS-LB <5% · IB1 5–9% · IB2 10–19%CD117+CD33+Aberrant CD5/CD7/CD56 on blasts
★ Ogata/Wells-Ogata FC score: validated quantitative MDS scoring. Score ≥2 = high sensitivity/specificity for MDS. MDS-h: BM ≤25% cellularity; T-cell immune attack; overlaps AA/PNH; IST-responsive subset
CD34 IHC — quantify blast %p53 IHC (if TP53 suspected)Reticulin stain (mandatory · MF grade 0–3)
★ Auer rod in ANY blast = MDS-IB2 regardless of blast % (confirm with MPO stain). ALIP = clusters of ≥3–5 CD34+/MPO+ blasts in central intertrabecular space = adverse marker. Reticulin grading mandatory in all MDS trephine reports
TET2 · DNMT3A · ASXL1 · SRSF2 · U2AF1 · ZRSR2 · RUNX1 · EZH2 · STAG2 · IPSS-R and IPSS-M risk stratification · HMA (azacitidine/decitabine) · Luspatercept (SF3B1-mutated) · Imetelstat (LR-MDS) · AlloSCT high/very-high risk
MDS/MPN Overlap
CMML (Types 1 & 2) · MD-CMML vs MP-CMMLMonocyte-committed progenitor / HSC Classical monocytes (CD14+CD16−) ≥94% of monocyte gate — most specific flow finding for CMMLCD14+CD64+CD163+CD56− (aberrant CD56 = worse prognosis)
★ WHO 5 lowers monocytosis threshold to ≥0.5×10⁹/L. ICC: same threshold + requires clonality (≥1 myeloid mut VAF ≥10%). Both WHO 5 and ICC eliminate CMML-0. MP-CMML (WBC ≥13) = RAS pathway mutations + worse prognosis
CD14+CD163+Lysozyme+CD34+ (increases with CMML type)
★ CMML-1: <5% PB / <10% BM blasts · CMML-2: 5–19% PB / 10–19% BM or Auer rods · Promonocytes = blast equivalents · monoblasts + promonocytes ≥20% = AML transformation
TET2 (~60%) · SRSF2 (~50%) · ASXL1 (~40%) · RAS pathway (NRAS/KRAS/CBL) · TET2+SRSF2 co-mutation highly specific for CMML · RUNX1 (adverse) · Ruxolitinib (WBC >13) · Azacitidine · AlloSCT high-risk
Acute Myeloid Leukaemia
APL with PML::RARA ⚠ EMERGENCYPromyelocyte-stage progenitor CD33+++ (very bright)CD13+CD34− (hypergranular)HLA-DR− (KEY absence)CD11b−CD2+ (microgranular M3v)
⚑ HLA-DR−/CD34− + coagulopathy = APL → START ATRA EMPIRICALLY. M3v mimics monocytic AML — CD34−/HLA-DR− pattern is the clue. Both WHO 5 and ICC: ≥10% blasts required
PML::RARA FISH+ (t(15;17))PML IHC: cytoplasmic diffuse pattern (normal = nuclear dots)MPO++ (very strong)
★ ATRA + ATO = chemo-free standard for standard-risk APL. High-risk (WBC >10): add GO. MRD by PML::RARA RT-PCR (IS) mandatory. ZBTB16::RARA (PLZF) = ATRA-resistant — always identify fusion partner
t(15;17)(q24.1;q21.2) · ATRA+ATO (standard-risk) · ATRA releases HDAC repression → differentiation · ATO degrades PML::RARA · MRD by RT-PCR IS at each treatment milestone
AML with RUNX1::RUNX1T1 · t(8;21)GMP / myeloid progenitor CD34+CD117+CD33+CD19+ (aberrant B-marker on myeloid blasts)CD56+ (subset — adverse)MPO+
★ CD19+ on myeloid blasts = classic t(8;21). Any CD19+ myeloid blast → RUNX1::RUNX1T1 FISH/RT-PCR urgently. WHO 5: any blast %; ICC: ≥10%
MPO+ (strong)CD34+CD19+
★ Salmon-pink granules + long single Auer rods + CD19+ aberrant = classic triad. RUNX1::RUNX1T1 can be cryptic on ~30% of standard karyotypes → FISH mandatory. MRD by RT-PCR IS
RUNX1::RUNX1T1 · KIT D816V/Y (~25%) adverse · NRAS/KRAS · HiDAC ×4 cycles consolidation · GO benefit in CBF-AML
AML with CBFB::MYH11 · inv(16)GMP / myeloid + monocytic + eosinophilic CD34+CD117+CD33+CD2+ (aberrant T-marker on myeloid blasts)MPO+CD64+
★ CD2+ on myeloid blasts = classic inv(16). CBFB FISH break-apart probe required — inv(16) frequently missed on G-banding
MPO+Abnormal eosinophils in BM: enlarged purple/basophilic granules in eosinophilic cells — essentially pathognomonic for inv(16)
★ Abnormal mixed-granule eosinophils (purple+eosinophilic granules in same cell) on BM aspirate = pathognomonic for inv(16)
CBFB::MYH11 · KIT exon 8 (~40%) · NRAS · HiDAC consolidation · GO benefit · MRD by RT-PCR IS
AML with NUP98 rearrangement — NEW in WHO 5 · NOT in ICCHSC / myeloid progenitor; variable differentiation CD34+CD117+Variable differentiation · NUP98::KDM5A = megakaryoblastic (CD41/CD61+)
⚑ NUP98 rearrangements CRYPTIC on standard karyotype — RNA-seq or NUP98 break-apart FISH required. WHO 5-specific entity: NOT in ICC 2022. Any pediatric/young adult AML with poor prognosis and no identified genetic driver should have NUP98 testing
MPO+ (most)CD34 variable
★ NUP98 has >30 fusion partners. Common: NUP98::NSD1 (pediatric/young adult poor prognosis), NUP98::HOXA9, NUP98::KDM5A (pediatric AMKL). All deregulate HOX gene expression. WHO 5: diagnosed at any blast %. This is a WHO 5-exclusive entity
NUP98 11p15.4 rearrangement · >30 partners · NUP98::NSD1 (poor prognosis, young adults) · NUP98::KDM5A (pediatric AMKL, CBFA2T3::GLIS2 spectrum) · NUP98::HOXA9 · HOX deregulation common mechanism · NOT in ICC 2022
AML with NPM1 mutation · ~30% of AMLGMP / mixed differentiation CD33+CD13+HLA-DR+CD34− (paradox — most NPM1-mutated AML is CD34 negative)CD14+/CD64+ if monocytic
★ CD34-negative AML with normal karyotype = NPM1 mutation until proven otherwise. CD34-negativity occurs because mutant NPM1c exported to cytoplasm. CD34-negative AML must automatically trigger NPM1 testing
NPM1 IHC: cytoplasmic staining (NPM1c+) — reliable surrogate on trephineCD34− (most)Cup-like nuclear invaginations on Romanowsky stain
★ Normal NPM1 = nuclear dots; mutant NPM1c = cytoplasmic diffuse. NPM1 IHC can be performed on decalcified trephine sections. Revumenib (menin inhibitor) Phase III AUGMENT-101 for NPM1-mutated AML
NPM1 exon 12 insertion (type A ~80%) · FLT3-ITD allelic ratio (AR): >0.5 = high AR = adverse · DNMT3A · IDH1/2 · TET2 · Midostaurin + 7+3 if FLT3+ · NPM1 ddPCR/RQ-PCR for MRD = gold standard
AML with CEBPA mutation (in-frame bZIP)GMP / granulocytic progenitor CD34+CD117+CD13+HLA-DR+CD7+ (aberrant T-marker)CD14−
⚑ WHO 5 requires ≥20% blasts · ICC requires ≥10% blasts for this entity — a key classification-dependent blast threshold difference. Both require in-frame bZIP (C-terminal) mutation
MPO+CD34+CD7+
★ WHO 5 updated: monoallelic bZIP also qualifies (favorable). ICC: bZIP mutations included (≥10% blasts). Both: in-frame bZIP domain required. Germline CEBPA: autosomal dominant AML predisposition — test all family members
CEBPA in-frame bZIP (C-terminal) · Mono- or bi-allelic bZIP both favorable (WHO 5) · Germline CEBPA risk · GATA2 co-mut · TET2/WT1 · Excellent prognosis (~60% 5-yr OS) · AlloSCT CR1 often deferred
AML with KMT2A rearrangement · 11q23.3HSC / myeloid-monocytic progenitor CD33+++ (characteristically very bright)CD64+CD14+ (monocytic)NG2 (clone 7.1)+ — highly specific for KMT2A-rearranged AML and ALLCD34±
★ CD33 characteristically very bright in KMT2A-r AML. NG2 (clone 7.1) = highly specific for KMT2A rearrangement in AML AND ALL contexts. Any blast % in both WHO 5 and ICC
NG2 (clone 7.1)+CD33+Lysozyme+
★ t(9;11) MLLT3::KMT2A = intermediate · t(6;11) AFDN::KMT2A = poor · t(10;11) = variable. Revumenib (menin inhibitor) FDA-approved Oct 2024 for r/r KMT2A-rearranged AML/ALL. Mixed-lineage surface markers (CD19, CD2) can occur — check MPAL criteria before reclassifying
KMT2A rearrangement 11q23.3 · >80 partners · Revumenib FDA 2024 · Ziftomenib (menin) Phase III · FLT3 co-mut frequent · AlloSCT CR1 recommended for most KMT2A-r AML
AML-MR (WHO 5 8 genes) · AML with MR gene mutations (ICC 9 genes)HSC; myelodysplastic evolution CD34+CD117+CD33+/CD13+Aberrant CD5/CD7/CD56 on blasts
⚑ WHO 5 AML-MR = 8-gene list (NO RUNX1): ASXL1, BCOR, EZH2, SF3B1, SRSF2, STAG2, U2AF1, ZRSR2. ICC = 9-gene list (adds RUNX1). This is a clinically important difference — affects CPX-351 eligibility. Both require ≥20% blasts for AML designation
CD34+p53 IHC (if TP53 co-mutation)
★ CPX-351 (liposomal daunorubicin+cytarabine) = standard induction for AML-MR (FDA-approved, superior to 7+3). WHO 5: ≥20% blasts required. ICC: 10–19% blasts = MDS/AML with MR gene mutations (different treatment implications)
WHO 5 AML-MR 8 genes: ASXL1, BCOR, EZH2, SF3B1, SRSF2, STAG2, U2AF1, ZRSR2 (NO RUNX1) · ICC adds RUNX1 = 9 genes · CPX-351 standard · Venetoclax+HMA unfit patients · AlloSCT eligible
BPDCNPlasmacytoid dendritic cell precursor CD123++ (exceptionally bright)CD4+CD56+HLA-DR+CD3−CD19−MPO−CD14−
★ CD123++ CD4+ CD56+ without other lineage markers = BPDCN. WHO 5 Table 15 adds CD303 (BDCA-2) and CD304 (BDCA-4) as expected positive pDC markers in updated criteria
TCF4 (E2-2) IHC+ — most specific BPDCN markerSPIB IHC+ — highly sensitiveCD123++CD4+
★ WHO 5 BPDCN diagnostic criteria (Table 15): CD123 + 1 other pDC marker (TCF4/TCL1/CD303/CD304) + CD4 and/or CD56 OR any 3 pDC markers + all negatives absent. Tagraxofusp-erzs (CD123 × diphtheria toxin) FDA 2018
TET2 (~50%) · NRAS/KRAS · ASXL1 · ZEB2 · MYC amplification · CDKN2A/B del · Tagraxofusp-erzs FDA 2018 · AlloSCT consolidation in CR1 · CAR-T (CD123) in trials

Purple border = hallmark/discriminating · Green = positive · Red-orange = negative · Amber = variable

EntityBM Biopsy ArchitectureAspirate / PB CytologyKey Diagnostic FeaturesPitfalls / Must Not Miss
MPN — Megakaryocyte Morphology (the Critical MPN Trephine Challenge)
CML Near 100% cellularity · Granulocytic hyperplasia overwhelming fat · Pseudo-Gaucher cells (lipid-laden macrophages) · Sea-blue histiocytes · No significant fibrosis in CP · WHO 5: only CP and BC. ICC: AP at 10–19% blasts Left-shifted granulocytosis · "Myelocyte bulge": myelocytes > metamyelocytes (inverse of reactive) · Basophilia (hallmark — virtually no other condition causes this degree) · Eosinophilia · NAP score very LOW (contrast reactive = HIGH)
  • Basophilia in PB = CML until proven otherwise
  • Myelocyte bulge distinguishes from reactive left shift
  • Pseudo-Gaucher cells = high granulocyte turnover marker
  • WHO 5: AP eliminated. ICC: AP = 10–19% blasts OR basophils ≥20% OR major route ACA
⚑ Leukemoid reaction vs CML: reactive lacks basophilia, has high NAP, BCR::ABL1 negative. Always test BCR::ABL1 for any unexplained WBC >25×10⁹/L
ET vs pre-PMF — Critical MPN Morphological Distinction ET: normocellular or mildly hypercellular · Megakaryocytic hyperplasia ONLY · Fat preserved · MF-0
pre-PMF: Hypercellular · Granulocytic + megakaryocytic hyperplasia · TIGHT dense mega clusters · MF-0/1 at most
Overt PMF: MF-2/3 · osteosclerosis · sinusoidal hematopoiesis · dry tap
ET: Thrombocytosis >450 · large/giant platelets · No left shift
pre-PMF: May look like ET early · Circulating CD34+ >10/μL (discriminating by flow)
Overt PMF: Leukoerythroblastic PB · teardrop cells · circulating megakaryocyte nuclear fragments
  • ET megakaryocytes: large, mature, hyperlobated (staghorn ≥5 lobes), loosely scattered or SMALL loose clusters (ICC: ≤6 cells)
  • pre-PMF megakaryocytes: TIGHT dense clusters (>6 cells ICC), bulbous/hyperchromatic nuclei, "naked nuclei" (extruded megakaryocyte segments) = HALLMARK
  • Reticulin staining (MF-0 to MF-3) is mandatory in ALL MPN trephine reports
  • Teardrop cells + leukoerythroblastosis = myelofibrosis until proven otherwise
⚑ pre-PMF vs ET = SINGLE most critical MPN morphological distinction. pre-PMF has higher MF transformation risk, different prognosis and therapy implications. Many pre-PMF cases are incorrectly called ET — requires expert trephine evaluation
Systemic mastocytosis Multifocal dense MC aggregates ≥15 MCs/cluster = major criterion · Paratrabecular or perivascular clustering · Fibrosis around aggregates · MCL: diffuse MC infiltrate ≥20% of nucleated BM cells · SM-AHN/SMN: concurrent AML/MDS/MPN morphology Spindle-shaped (fusiform) mast cells on BM aspirate (≥25% = minor criterion) · Normal MCs = round/oval; neoplastic = spindle · Metachromatic granules (Giemsa/toluidine blue positive) · Eosinophilia in PB (PDGFRA-associated)
  • Spindle-shaped MCs = neoplastic (normal = round/oval)
  • Tryptase IHC highlights aggregates · CD25 IHC confirms aberrant expression
  • WDSM: round MCs · KIT D816V absent · CD25−/CD30+ · fails standard minor criteria
  • B-findings (burden): BM >30% MC, tryptase >200, organomegaly, dysplasia without cytopenia
  • C-findings (damage): cytopenias, liver/spleen/gut dysfunction, bone fractures
⚑ MC aggregates mimicked by fibroblasts or plasma cell clusters on H&E — tryptase IHC + CD25 IHC mandatory for confirmation before diagnosing SM
MDS-5q Variable cellularity · Megakaryocytic hyperplasia · Small MONOLOBATE megakaryocytes (pathognomonic) · Erythroid hypoplasia + dysplasia · Granulopoiesis preserved Macrocytic anemia · Thrombocytosis or normal platelet count · Normal or mildly elevated WBC · Rare circulating blasts
  • Monolobate megakaryocytes on CD61 IHC = histological hallmark of del(5q)
  • Normal: ≥3 nuclear lobes; del(5q): predominantly 1 lobe
  • FISH confirmation mandatory before lenalidomide · TP53 sequencing before prescribing
⚑ Monolobate megakaryocytes + macrocytic anemia in elderly woman = del(5q) until proven otherwise. Additional del(7q)/−7 changes entity — review full karyotype before lenalidomide
MDS-LB / MDS-IB1 / MDS-IB2 / MDS-F / MDS-h Variable cellularity · Dysplasia ≥10% in ≥1 lineage · ALIP (central intertrabecular CD34+/MPO+ clusters) · MDS-F: MF-2/3 · MDS-h: ≤25% cellularity + clonal dysplasia Cytopenias · Pseudo-Pelger-Huët cells (bilobed, hypogranular) · Hypogranular neutrophils · MDS-IB: circulating blasts · Auer rods (→ always MDS-IB2 regardless of blast %) · Dimorphic RBCs
  • Dysplasia: ≥10% of cells in a lineage (both WHO 5 + ICC)
  • Auer rod in ANY blast = MDS-IB2 regardless of blast % (confirm with MPO stain)
  • ALIP: ≥3–5 CD34+/MPO+ cells in central intertrabecular space = adverse prognostic marker
  • Perl's Prussian blue stain MANDATORY in all MDS workups
  • MDS-h: BM ≤25% cellularity + dysplasia + clonality · overlaps AA/PNH; germline testing essential
⚑ Dysplasia on aspirate can be artifactual (poor smear quality, prolonged EDTA) — always assess smear quality before counting. Acquired pseudo-Pelger vs inherited Pelger-Huët anomaly: inherited = bilobate but normally granulated + no other cytopenias
CMML Variable cellularity · Monocytic expansion (interstitial CD68/CD163+) · Granulocytic dysplasia · CD34+ blasts increase with type · Fibrosis in subset Monocytosis ≥0.5×10⁹/L AND ≥10% WBC · Promonocytes (blast equivalents) = large cells with irregular nuclear folds, fine chromatin, pale gray cytoplasm · Dysplastic neutrophils · Proliferative CMML: WBC ≥13
  • Promonocytes = blast equivalents; monoblasts + promonocytes ≥20% = AML transformation
  • CMML-1: <5% PB / <10% BM · CMML-2: 5–19% PB / 10–19% BM or Auer rods
  • Classical monocytes ≥94% on flow partitioning = CMML vs reactive monocytosis
  • ICC: also requires clonality evidence in most cases
⚑ Promonocytes vs monocytes: promonocytes have irregular nuclear folds, fine chromatin, ground-glass cytoplasm — distinct from mature kidney-shaped monocyte nuclei. Under-counting promonocytes underestimates blast equivalents
APL with PML::RARA ⚠ MEDICAL EMERGENCY Hypercellular BM · Sheets of abnormal promyelocytes · Hypergranular: coarse granules obscuring nuclear outline · M3v: bilobed nuclei + fine dusty granules Hypergranular promyelocytes (coarse azurophilic granules, bilobed "butterfly" nuclei) · FAGGOT CELLS: bundles of multiple Auer rods per cell = PATHOGNOMONIC · DIC evidence: schistocytes + thrombocytopenia · M3v: bilobed cells with fine granules — mimics monocytic AML
  • Faggot cells (bundles of Auer rods per cell) = APL morphological EMERGENCY
  • DIC in any new AML = APL until PML::RARA FISH result available
  • ATRA EMPIRICALLY while FISH pending — NEVER wait
  • Both WHO 5 and ICC: ≥10% blasts required for APL
  • Distinguish: single long Auer rod (t(8;21)) vs bundled Auer rods (Faggot cells, APL)
⚑ MOST DANGEROUS morphological pitfall in hematopathology: missing APL = fatal DIC. ANY highly granular myeloid blast + coagulopathy → START ATRA IMMEDIATELY
⚑ M3v mimics monocytic AML — CD34−/HLA-DR− on flow + coagulopathy = APL regardless of granule appearance
AML with RUNX1::RUNX1T1 · t(8;21) BM replaced by blasts · Large blasts with abundant cytoplasm · Salmon-pink primary granules · Long single Auer rods prominent · Residual dysplastic granulocytes Salmon-pink cytoplasm + long single Auer rods (solitary per cell) + CD19+ aberrant on myeloid blasts = CLASSIC TRIAD
  • Salmon-pink abundant cytoplasm + long single Auer rods + CD19 coexpression = t(8;21) presumptive
  • Can be diagnosed at <20% blasts (any blast % in WHO 5; ≥10% in ICC)
  • RUNX1::RUNX1T1 can be cryptic (~30% standard karyotypes) → FISH mandatory
⚑ Do NOT mistake long single Auer rods for Faggot cells (APL). Single rod per cell = t(8;21); bundled multiple rods per cell = APL. If any doubt, rule out APL first
AML with CBFB::MYH11 · inv(16) Hypercellular BM · Myeloid + monocytic + eosinophilic components Abnormal eosinophils = PATHOGNOMONIC: immature cells containing BOTH eosinophilic AND enlarged purple/basophilic granules in same cell
  • Abnormal mixed-granule eosinophils (purple+eosinophilic) essentially pathognomonic for inv(16)
  • CD2+ on myeloid blasts + abnormal BM eosinophils = order CBFB FISH urgently
  • inv(16) frequently missed on G-banding → CBFB break-apart FISH mandatory
⚑ inv(16) is the cytogenetic finding most commonly missed on standard G-banding. FISH with CBFB break-apart probe is mandatory in ALL myelomonocytic AML with eosinophilia regardless of karyotype result
AML with NUP98 rearrangement (WHO 5 ONLY entity) Variable cellularity · Blast infiltration · No specific BM architectural pattern Variable cytology by fusion partner · NUP98::KDM5A = megakaryoblastic (CD41/CD61+) · NUP98::HOXA9 = poorly differentiated · No pathognomonic cytomorphological feature
  • No pathognomonic morphological feature — molecular confirmation required
  • CRYPTIC on standard karyotype — RNA-seq or NUP98 FISH required
  • WHO 5-only entity: NOT in ICC 2022 — institutional awareness needed to test for it
⚑ NUP98 rearrangements will be missed without RNA sequencing or targeted FISH. Any pediatric/young adult AML with normal karyotype + poor response + no other identified genetic driver should have NUP98 testing
AML with NPM1 mutation Hypercellular BM · Usually normal karyotype · Often monocytic/myelomonocytic differentiation · CD34-negative blast clusters on IHC (unusual) Cup-like nuclear invaginations (punched-out nuclear indentation) in blasts = characteristic NPM1 AML feature on Romanowsky stain · Monocytic differentiation common · Occasional Auer rods
  • Cup-like nuclear morphology in blasts = characteristic NPM1 AML clue
  • CD34-negative blast clusters on IHC = unusual → NPM1 testing mandatory
  • NPM1 IHC cytoplasmic staining on trephine = rapid surrogate for NPM1 mutation
⚑ CD34-negative AML should AUTOMATICALLY trigger NPM1 mutation testing — this is standard of care
⚑ Always test FLT3 allelic ratio concurrently with NPM1 — allelic ratio (>0.5 high; <0.5 low) determines prognosis and guides alloSCT decision in CR1

⚑ = diagnostic pitfall · ★ = high-yield discriminating feature · ⚠ APL = emergency — ATRA empirically while FISH pending

Legend: Concordant = essentially same criteria · Minor divergence = naming or minor criteria differences · MAJOR divergence = different entities, thresholds, or clinical implications
TopicWHO 5 (Khoury et al., Leukemia 2022)ICC 2022 (Arber et al., Blood 2022)Divergence LevelClinical Impact / Pitfall
CML — Phase Classification
CML phases WHO 5: Eliminates accelerated phase (AP). Only chronic phase (CP) and blast phase (BP). Emphasis on risk features in CP. BP: ≥20% blasts OR extramedullary blasts OR ↑ lymphoblasts in PB/BM ICC: Retains accelerated phase (AP). AP = blasts 10–19% OR PB basophils ≥20% OR major route ACA in Ph+ cells (second Ph, +8, i17q, +19, complex, abn 3q26.2) MAJOR ⚑ WHO 5: A patient with 15% blasts in CML = still "chronic phase with high-risk features" + TKI optimization. ICC: same patient = accelerated phase, may qualify for different trial eligibility or alloSCT consideration. Report both classifications in pathology report when divergence matters clinically
JMML Categorization
JMML classification placement WHO 5: JMML is placed under MPN. Reflects RAS pathway molecular pathogenesis and absence of bona fide MDS stigmata. KMT2A rearrangements excluded from JMML diagnosis in WHO 5 ICC: JMML placed under Pediatric and/or germline mutation-associated disorders. ICC also defines JMML-like neoplasms (no RAS mutation) and Noonan syndrome-associated MPD as formal entities Minor ⚑ Placement differs between WHO 5 (MPN) and ICC (Pediatric). Clinical criteria are essentially the same but ICC provides additional guidance on JMML-like neoplasms and Noonan-MPD which WHO 5 doesn't formalize as separate entities. ICC Table 22 requires monocytes ≥1×10⁹/L (not ≥0.5 like CMML); ~7% JMML cases may not meet this criterion
AML — NUP98 Rearrangement
AML with NUP98 rearrangement WHO 5 Table 7: NEW standalone AML entity. Any blast % (KMT2A/MECOM/NUP98 are the three rearrangement-based entities that allow any blast %). WHO 5 explicitly states NUP98 rearrangements may be cryptic on conventional karyotyping ICC 2022: NUP98 rearrangement is NOT a separate AML entity. Cases with NUP98 fusions would fall into AML with other MDS-related cytogenetic abnormalities, AML-NOS, or possibly MDS/AML depending on blast count MAJOR ⚑ Critical practical pitfall: A pediatric patient with 12% blasts and NUP98::NSD1 fusion = "AML with NUP98 rearrangement" (WHO 5, any blast %) vs "MDS/AML with MR cytogenetics or MDS-IB2" (ICC). Treatment intensity differs dramatically. Institutions using ICC must have an institutional protocol to test for and flag NUP98 rearrangements given their clinical importance
AML — TP53 Mutated AML
AML with mutated TP53 WHO 5: AML with mutated TP53 is NOT a standalone separate AML entity. Biallelic TP53 inactivation is recognized as MDS-biTP53 (in MDS) and within AML-MR (cytogenetic criteria). WHO 5 does not create a standalone "AML with mutated TP53" category ICC 2022 (Table 21): Creates a standalone TP53-mutated spectrum category. MDS with mutated TP53 (multi-hit required) · MDS/AML with mutated TP53 (10–19% blasts, any TP53 VAF >10%) · AML with mutated TP53 (≥20% blasts, any TP53 VAF >10%). Groups these together due to similar aggressive biology regardless of blast % MAJOR ⚑ ICC AML with mutated TP53 = any pathogenic TP53 mutation VAF >10% (monoallelic allowed for AML). WHO 5 handles TP53-mutated AML within AML-MR cytogenetic category. This ICC-specific framing groups MDS, MDS/AML, and AML with TP53 together for unified treatment strategy. Pure erythroid leukemia in ICC is classified within AML with mutated TP53 (TP53 mutations very common in AEL)
AML — CEBPA Blast Threshold
AML with CEBPA mutation — blast threshold WHO 5: Requires ≥20% blasts (exception along with BCR::ABL1). "There is insufficient data to support any change in the blast cutoff criterion for AML with CEBPA mutation" (Khoury et al., 2022). In-frame bZIP; monoallelic bZIP now qualifies ICC 2022 (Table 25): Requires ≥10% blasts (same threshold as most other genetic AML entities). ICC also requires in-frame bZIP mutations. Unlike WHO 5, ICC lowers the blast threshold for CEBPA MAJOR ⚑ A patient with 15% blasts and monoallelic bZIP CEBPA mutation: WHO 5 = NOT AML with CEBPA (only MDS-IB2 since <20%). ICC = AML with in-frame bZIP CEBPA mutations (favorable prognosis entity). This directly affects whether a patient receives HMA (MDS) vs intensive induction chemotherapy. Additionally, WHO 5 updated to include monoallelic bZIP as qualifying (equivalent prognosis to biallelic bZIP-containing cases). Both WHO 5 and ICC now agree that monoallelic in-frame bZIP mutations qualify for the favorable entity
AML-MR / AML-MRC — Gene List
Myelodysplasia-related gene mutation list WHO 5 AML-MR (Table 8): 8 somatic genes — ASXL1, BCOR, EZH2, SF3B1, SRSF2, STAG2, U2AF1, ZRSR2. RUNX1 is NOT on the WHO 5 AML-MR list (WHO 5 explicitly states: "RUNX1 mutations in AML overlap with such a broad range of defining molecular features that it was determined to lack enough specificity to define a standalone AML type") ICC 2022 (Table 25): 9 somatic genes — ASXL1, BCOR, EZH2, RUNX1, SF3B1, SRSF2, STAG2, U2AF1, ZRSR2. RUNX1 IS on the ICC list (now encompasses the prior provisional entity of "AML with mutated RUNX1"). Blast threshold: ≥20% for AML; 10–19% = MDS/AML MAJOR ⚑ A patient with ≥20% blasts and a RUNX1 mutation only (no other MR feature): WHO 5 = AML-NOS (not AML-MR since RUNX1 not on WHO 5 list). ICC = AML with myelodysplasia-related gene mutations (eligible for CPX-351). A patient with 15% blasts and SRSF2 mutation: WHO 5 = MDS-IB2 (HMA standard). ICC = MDS/AML with MR gene mutations (potentially CPX-351 eligible). These are clinically high-stakes differences for treatment decisions
MDS — MDS/AML Bridging Category and Blast Thresholds
MDS-IB2 vs MDS/AML (10–19% blasts) WHO 5: 10–19% BM blasts without defining genetic lesion = MDS-IB2 (not AML). WHO 5 retains ≥20% blast threshold for MDS → AML transition. "Broad agreement that MDS-IB2 may be regarded as AML-equivalent for therapeutic considerations" (Khoury et al., 2022) ICC 2022: Introduces MDS/AML as a new category for 10–19% blasts (adults; pediatric cases retain MDS-EB with 10–19% blasts). Eligible for both MDS and AML clinical trials. Sub-classified as MDS/AML-TP53, MDS/AML-MR gene muts, MDS/AML-MR cytogenetics, MDS/AML-NOS MAJOR ⚑ Most clinically important classification divergence for treatment decisions. At 15% blasts with SRSF2 mutation: WHO 5 = MDS-IB2 (HMA standard, may consider CPX-351 off-label). ICC = MDS/AML with MR gene mutations (CPX-351 appropriate, eligible for AML trials). Approved drug labeling (CPX-351, venetoclax) uses ≥20% blasts — the ICC MDS/AML framing currently exceeds regulatory approvals, creating a treatment gap
MDS — SLD vs MLD Distinction
Single vs multilineage dysplasia WHO 5: MDS-LB merges single lineage dysplasia (SLD) and multilineage dysplasia (MLD) into one category. "The distinction between single lineage and multilineage dysplasia is now considered optional" (Khoury et al., 2022) ICC 2022: Retains SLD and MLD as separate subtypes within MDS-NOS. "Although there is poor reproducibility in distinguishing single lineage vs multilineage dysplasia in MDS, this distinction has been retained" (Arber et al., 2022) Minor ⚑ WHO 5 MDS-LB does not distinguish SLD from MLD — loses prognostic granularity used in clinical trials and some registry databases. Institutions using WHO 5 that report only "MDS-LB" cannot satisfy trial eligibility criteria that specify SLD vs MLD. Consider documenting number of dysplastic lineages even when using WHO 5 MDS-LB designation
MDS/MPN — aCML Naming
aCML / MDS/MPN with neutrophilia WHO 5: Renames atypical CML to "MDS/MPN with neutrophilia" (MDS/MPN-N). The name "atypical CML" is formally retired to avoid confusion with CML ICC 2022: Retains "atypical CML, BCR::ABL1-negative" as the preferred name. Drops the "BCR::ABL1-negative" qualifier but keeps "aCML". Same diagnostic criteria Minor — naming only ⚑ Same entity, same criteria, different names. Pathology reports using "aCML" (ICC) vs "MDS/MPN-N" (WHO 5) for the same case may appear as different diagnoses in registries. Reports should ideally cross-reference both terms when the two classifications assign different names
MDS/MPN — ICC-only entities
CMUS / CCMUS · MDS/MPN with i(17q) WHO 5: CCUS is defined (CHIP + unexplained cytopenia) but CMUS (with monocytosis) is not a distinct WHO 5 entity. WHO 5 acknowledges "additional studies needed to determine optimal approach to classifying individuals with unexplained clonal monocytosis who do not fit the new CMML diagnostic criteria." MDS/MPN with i(17q) not listed as provisional entity in WHO 5 ICC 2022: CMUS (clonal monocytosis of undetermined significance) = formal ICC entity for cases with monocytosis ≥0.5×10⁹/L + clonality but NO BM findings of CMML. CCMUS if cytopenia also present. MDS/MPN with i(17q) = new provisional subentity under MDS/MPN-NOS Minor — ICC provides more operational guidance ⚑ ICC CMUS/CCMUS provides explicit guidance for a common diagnostic grey zone (monocytosis + clonal mutations + no CMML morphology). WHO 5 acknowledges the grey zone but doesn't formalize a diagnostic category. In practice, ICC's CMUS concept is clinically useful and many centers apply it even when using WHO 5 terminology
SM — AHN vs AMN Terminology
SM-AHN (WHO 5) vs SM-AMN (ICC) WHO 5: Retains SM-AHN (SM with associated haematological neoplasm) — "haematological" allows for both myeloid AND lymphoid associated neoplasms. SM can have associated lymphoid neoplasm in WHO 5 ICC 2022: Renames to SM-AMN (SM with associated myeloid neoplasm). ICC restricts the associated neoplasm to myeloid context only, based on evidence that shared KIT mutations are present with myeloid but not lymphoid neoplasms Minor ⚑ Rare but important: SM + concurrent lymphoma (e.g., CLL/SLL) = "SM-AHN" in WHO 5 (formally recognized as an SM subtype) but would NOT be SM-AMN in ICC (which restricts to myeloid neoplasms). Clinically uncommon but the name matters for registry coding
Histiocytic/DC Neoplasms — FDCS Placement
Follicular dendritic cell sarcoma (FDCS) placement WHO 5: FDCS and fibroblastic reticular cell tumour are moved OUT of histiocytic/DC neoplasms to a new chapter: "stroma-derived neoplasms of lymphoid tissues" ICC 2022: Changes to acute leukemias of ambiguous lineage not addressed in the CAC meeting. ICC does not specifically address FDCS placement (lymphoid neoplasms handled separately) Minor — WHO 5 reorganization ⚑ Practical impact: FDCS should no longer be coded/filed under histiocytic neoplasms in WHO 5-based laboratory information systems. Reclassification from "histiocytic/DC" to "stroma-derived" has registry and coding implications
Summary: Where Each Classification Misses the Mark
WHO 5 — areas of concern
  • Loss of SLD/MLD distinction in MDS-LB: collapsing to one category loses prognostic information used by clinicians and trial designers
  • CEBPA blast threshold (≥20%): requiring 20% blasts for CEBPA AML means patients with 10–19% blasts and favorable bZIP mutations are classified as MDS-IB2 and may not receive appropriate intensive therapy
  • No CMUS/CCMUS concept: leaves a common diagnostic grey zone (monocytosis + clonality + no CMML morphology) without formal guidance
  • No standalone TP53-mutated AML entity: monoallelic TP53 AML biology is well-recognized as aggressive; WHO 5 does not create a distinct category for it, unlike ICC
  • NUP98 rearrangement entity vs ICC: institutions using ICC will miss this entity designation without actively cross-referencing WHO 5
ICC 2022 — areas of concern
  • MDS/AML category (10–19% blasts) lacks regulatory drug approvals: CPX-351, venetoclax approvals are for ≥20% blasts. The ICC MDS/AML framing is intellectually coherent but currently exceeds drug approval thresholds, creating a treatment ambiguity gap
  • BCR::ABL1 AML not a separate entity: ICC's framing of BCR::ABL1+ acute leukemia as CML blast crisis may route truly de novo BCR::ABL1+ AML patients to CML-phase treatment algorithms, potentially suboptimally
  • RUNX1 on the MR gene list: RUNX1 mutations occur broadly across many AML/MDS contexts. Including RUNX1 as a standalone AML-MR qualifier (as ICC does) may over-classify some cases as AML-MR and drive CPX-351 use in situations where it may not be warranted
  • No NUP98 entity: NUP98 rearrangements are established as prognostically and therapeutically important; their absence from ICC leaves a gap that WHO 5 addresses
  • Retained legacy terminology (aCML, RCC, MDS-EB): keeping multiple names for the same entities increases cross-classification confusion for clinicians who move between WHO 5 and ICC institutions

WHO 5th Ed = Khoury JD, Solary E, Hochhaus A et al. Leukemia 2022;36:1703–1719 · ICC 2022 = Arber DA, Orazi A, Hasserjian RP et al. Blood 2022;140:1200–1228. Both effective simultaneously. Pathology reports should document which classification was applied. Clinical trials specify which classification was used for eligibility determination.